Having spent some of the time travelling and trekking in nearby hills and mountains i witnessed neps of same species often show slight variation in shapes and colors in different locality. An example N. edwardsiana found on Mt.Trus Madi are slightly different in shape from those found on Mt.Kinabalu,Lowii in Mt Trus Madi (2,462m)also differentiate in colors and shape from Mt.Murud(2,650m)
The lowland temperature and humidity is different as one move inlands and the differences will be greater at night.It will be even greater around hills and mountains as clouds descent due to condensation.Thus providing all the plants with the requirements.Inland tends to get more rain than places nearer the coast,as least in places around Borneo.
Having said that if we can trace the origin of the clone's parentage fine tuning their growth requirement as per their native habitats will enhance their colors and shapes. Post you comment.
Posted here 2 images - inland/lowland hills landscape.It was foggy at first but as the sun rises,the warm dissipate the fogs.This is a typical northiana's habitat,Bau limestone hill.
A typical highland scene in the morning. One of veitchii's habitats,Borneo highlands.The low clouds formation camouflage the hills and mountain.